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The North American Species of Cactus； Anhalonium； and Lophophora
by John M。 Coulter
A Preliminary Revision of the North American Species of Cactus； Anhalonium； and Lophophora by John M。 Coulter。
U。 S。 Department of Agriculture Division of Botany CONTRIBUTIONS FROM THE U。 S。 NATIONAL HERBARIUM Vol。 IIINo。 2 Issued June 10； 1894 Preliminary Revision of the North American Species of Cactus； Anhalonium； and Lophophora。 by John M。 Coulter。 Published by Authority of the Secretary of Agriculture Washington Government Printing Office 1894 LETTER OF TRANSMITTAL U。 S。 Department of Agriculture Division of Botany Washington； D。 C。； March 21； 1894 SIR： I have the honor to transmit herewith； for publication as Vol。 III； No。 2； of Contributions from the U。 S。 National Herbarium； a Preliminary Revision of the North American species of Cactus； Anhalonium； and Lophophora； by President John M。 Coulter。 Respectfully； Frederick V。 Coville； Chief of the Division of Botany。 Hon。 J。 Sterling Morton； Secretary of Agriculture。
PRELIMINARY REVISION OF THE NORTH AMERICAN SPECIES OF CACTUS； ANHALONIUM； AND LOPHOPHORA。 Prefatory Note。 In the fall of 1890 Dr。 George Vasey； then Botanist of the Department of Agriculture； arranged with me to prepare a revision of North American Cactaceae。 Owing to the peculiar difficulty of preserving material the family was poorly represented； even in our leading herbaria。 To secure a large amount of additional material in the way of specimens and field notes the Department authorized me to visit the region of the Mexican boundary during the summer of 1891。 Preliminary to this exploration it was necessary to examine the Engelmann collection of Cactaceae； in the possession of the Missouri Botanical Garden。 This collection； supplemented by the continual additions made at the garden； is by far the largest collection of skeletons and living specimens in this country； and also contains the large majority of our types。
In March； 1891； I visited this collection and made such notes as seemed necessary for use in the field； and in June； accompanied by Mr。 W。 H。 Evans and Mr。 G。 C。 Nealley； I began field work in the neighborhood of El Paso； Tex。 After ten days of exploration it was necessary for me to leave the field work in charge of Mr。 Evans； who； with Mr。 Nealley； continued work westward； during July and a part of August； to southern California； along the Southern Pacific Railway。 As a result a large number of complete plant bodies was secured； but very few of them were in flower and the field notes indicated little besides collection stations。 During the following fall and winter preliminary determinations of this material were made by Mr。 Evans。 In the fall of 1892 critical study of this and other collections was begun in connection with my assistants； Dr。 Elmon M。 Fisher and Mr。 Edwin B。 Uline； who have ever since rendered constant and most import assistance in the examination of material and bibliography； which alone has made the work possible in the midst of other pressing duties。
In the spring of 1893 these two gentlemen spent several weeks at the Missouri Botanical Garden in the critical study of its rich material； and during the latter part of their stay I assisted in the work。 Dr。 William Trelease； the director of the garden； had hastened the arrangement of the Engelmann material； and had mounted in convenient form the large mass of notes left by Dr。 Engelmann。 These notes contained not only critical remarks upon known species； but also the diagnoses of many unpublished species which had come into his hands； notably those collected by Mr。 William Gabb in 1867 in Lower California。 The collections that have thus far been studied are：
（1） Those of the Missouri Botanical Garden； and thanks are especially due to Dr。 Trelease for his generous cooperation in the use of this material； without which the work would have been impossible。
（2） Those of the Department of Agriculture； including the results of several recent explorations； for the use of which I am indebted to Mr。 Frederick V。 Coville。
（3） Those of the Gray Herbarium at Harvard University； which Dr。 B。 L。 Robinson kindly placed at my disposal。
（4） Those of the California Academy of Sciences； notably rich in forms from Lower California and the adjacent islands； kindly loaned by Mr。 T。 S。 Brandegee。
（5） Those of Dr。 Louis Eschanzier； of San Luis Potosi； Mexico； who send a large series of Mexican forms collected in 1891。
（6） Numerous small sets from different correspondents； who have given both time and material in aiding the work。
It is needless to say that Dr。 George Engelmann； the great pioneer student of this difficult family； has opened the paths in which we must follow； and it was exceedingly unfortunate that he was not able to complete the final revision that he had in mind。
The difficulties which beset the critical study of this group can not be easily exaggerated。 Such scanty material as has been collected has been for the most part very incomplete； consisting of plant bodies without flower or fruit； flower or fruit without plant bodies； and bunches of spines without either。 The species are displayed also in the most inaccessible regions； and their culmination is found in the still poorly known regions of Mexico。
On account of their singular forms and often brilliant flowers they have long been extensively cultivated； especially in Europe。 These cultivated forms have formed the basis of original descriptions in almost all of the European publications； and in very rare cases have any types been preserved。 As a result； the bibliography of Cactaceae is appalling； and it is questionable whether satisfactory conclusions can be reached in the case of hundreds of published names。 The earlier descriptions were not only meager； but were based upon what are now regarded very insufficient characters； and in the absence of types it is not only unsafe； but impossible to venture an opinion concerning their identity。 In view of these facts； I have thought it advisable to present a preliminary revision of the order； which shall contain the results of the study of material confessedly insufficient。 With such knowledge as we possess brought together； it is hoped that the study of this very interesting and much neglected group will be stimulated； and that more critical exploration of our southwestern territory and adjacent Mexico will make a more satisfactory presentation possible。 It would be useless to notice the vast number of reputed species that are not represented by actual specimens in our possession。
In the proposed preliminary account of the family； of which the present paper is the first part； only those genera are considered which form a part of the flora of the United States； and those species which I have been able to examine and to identify with reasonable certainty。 All forms credited to the United States have been studied； and the account of these species may be considered fairly complete； but the far more numerous Mexican species are but scantily represented。 The Mexican boundary is so unnatural a dividing line in the distribution of Cactaceae that it has been disregarded； and all the species studied have been arranged in a lineal series of uniform prominence。 So far as known the subject of geographical distribution is considered； but it will be seen how meager is our knowledge of this subject。 It is to be hoped that this preliminary presentation will provoke exploration and study； and that species will not only be collected； but all the facts of their distribution noted。 It is more than probable that our present notion of species in this group must be much modified； and doubtless many forms are at present kept specifically distinct which will prove to be but different phases of a single species。
In the matter of generic delimitation we are in still greater uncertainty； and several generic lines at present recognized must be regarded as purely arbitrary； a fact which must become still more evident with additional material。 The whole group is to be regarded as made up of poorly differentiated forms and only long observation under cultivation can determine the possibilities of specific variation under the influence of environment； of age； of inherent tendencies。 For instance； that these plants change in form and in spine characters with increasing age and after they have begun to flower can not be doubted； but what described forms have thus been separated in descriptions can only be guessed at。
John M。 Coulter。 Lake Forest University； Lake Forest； Ill。； January； 1891。
CACTUS； ANHALONIUM； AND LOPHOPHORA。
1。 CACTUS Linn。 Sp。 Pl。 466 （1753）； restricted。
MAMILLARIA Haw。 Synop。 177 （1812）； not Stackh。 （1809）。
Usually globose to oblong plants （simple； branching or cespitose）； but sometimes slender…cylindrical； covered with spine…bearing tubercles： flower…bearing areola axillary （with reference to tubercles）； entirely separate from the terminal spine…bearing areola； although sometimes （Coryphantha） connected with it by a woolly groove along the upper face of the tubercle： ovary naked： seeds smooth or pitted： embryo usually straight； with short cotyledons。 Orig